Plants of the Four Corners

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  Plantae
      Tracheophyta
            Magnoliopsida
                    Caryophyllales
                            Cactaceae
                              Cactoideae
                                     Acanthocalycium
                                             Acanthocalycium thionanthum
                                               Acanthocalycium thionanthum subsp. ferrarii
                                                     [Acanthocalycium ferrarii]
                                     Acharagma
                                             Acharagma roseanum
                                               Acharagma roseanum subsp. roseanum
                                                     [Escobaria roseana]
                                     Ariocarpus
                                             Ariocarpus fissuratus
                                                   [Mammillaria fissurata]
                                             Ariocarpus retusus
                                             Ariocarpus scaphirostris
                                     Astrophytum
                                             Astrophytum asterias
                                                   [Echinocactus asterias]
                                             Astrophytum capricorne
                                             Astrophytum myriostigma
                                             Astrophytum ornatum
                                     Bisnaga
                                             Bisnaga hamatacantha
                                               Bisnaga hamatacantha subsp. hamatacantha
                                                     [Echinocactus longihamatus]
                                                     [Ferocactus hamatacanthus]
                                     Borzicactus
                                             Borzicactus aureispinus
                                                   [Cleistocactus winteri]
                                             Borzicactus strausii
                                                   [Cleistocactus strausii]
                                     Carnegiea
                                             Carnegiea gigantea
                                                   [Cereus giganteus]
                                     Cephalocereus
                                             Cephalocereus senilis
                                     Cereus
                                             Cereus hildmannianus
                                               Cereus hildmannianus subsp. hildmannianus
                                                     [Cereus childsii]
                                             Cereus repandus
                                                   [Cereus peruvianus]
                                     Chamaecereus
                                             Chamaecereus silvestrii
                                                   [Echinopsis chamaecereus]
                                     Cleistocactus
                                             Cleistocactus smaragdiflorus
                                     Cochemiea
                                             Cochemiea barbata
                                                   [Mammillaria barbata]
                                             Cochemiea conoidea
                                                   [Echinocactus conoideus]
                                             Cochemiea dioica
                                                   [Mammillaria dioica var. incerta]
                                                   [Mammillaria dioica]
                                             Cochemiea grahamii
                                                   [Cactus grahamii]
                                                   [Mammillaria grahamii var. oliviae]
                                                   [Mammillaria grahamii]
                                                   [Mammillaria microcarpa var. auricarpa]
                                                   [Mammillaria microcarpa]
                                                   [Mammillaria milleri]
                                                   [Mammillaria oliviae]
                                                   [Neomammillaria microcarpa]
                                                   [Neomammillaria milleri]
                                                   [Neomammillaria oliviae]
                                             Cochemiea halei
                                                   [Mammillaria halei]
                                             Cochemiea mainiae
                                                   [Mammillaria mainiae]
                                             Cochemiea pondii
                                                   [Mammillaria pondii]
                                             Cochemiea setispina
                                                   [Mammillaria pondii subsp. setispina]
                                             Cochemiea tetrancistra
                                                   [Mammillaria tetrancistra]
                                             Cochemiea thornberi
                                               Cochemiea thornberi subsp. thornberi
                                                     [Mammillaria fasciculata]
                                                     [Mammillaria thornberi]
                                             Cochemiea viridiflora
                                                   [Mammillaria orestera]
                                                   [Mammillaria viridiflora]
                                             Cochemiea wrightii
                                               Cochemiea wrightii subsp. wilcoxii
                                                     [Mammillaria wilcoxii]
                                                     [Mammillaria wrightii subsp. wilcoxii]
                                                     [Mammillaria wrightii var. wilcoxii]
                                               Cochemiea wrightii subsp. wrightii
                                                     [Mammillaria meridiorosei]
                                                     [Mammillaria wrightii]
                                                     [Neomammillaria wrightii]
                                     Copiapoa
                                             Copiapoa grandiflora
                                             Copiapoa humilis
                                                   [Copiapoa humilis var. paposoensis]
                                             Copiapoa hypogaea
                                               Copiapoa hypogaea var. montana
                                                     [Copiapoa mollicula]
                                             Copiapoa krainziana
                                     Corryocactus
                                           [Erdisia]
                                     Coryphantha
                                             Coryphantha clavata
                                             Coryphantha compacta
                                             Coryphantha cornifera
                                                   [Coryphantha bernalensis]
                                                   [Coryphantha maliterrarum]
                                                   [Coryphantha radians]
                                                   [Coryphantha scolymoides]
                                             Coryphantha delaetiana
                                             Coryphantha delicata
                                                   [Coryphantha palmeri]
                                             Coryphantha durangensis
                                               Coryphantha durangensis subsp. cuencamensis
                                                     [Coryphantha cuencamensis]
                                             Coryphantha echinoidea
                                             Coryphantha echinus
                                                   [Coryphantha pectinata]
                                                   [Mammillaria echinus]
                                             Coryphantha elephantidens
                                               Coryphantha elephantidens subsp. bumamma
                                                     [Coryphantha bumamma]
                                               Coryphantha elephantidens subsp. elephantidens
                                                     [Coryphantha sulcolanata]
                                             Coryphantha erecta
                                             Coryphantha georgii
                                                   [Coryphantha grata]
                                                   [Coryphantha villarensis]
                                             Coryphantha maiz-tablasensis
                                             Coryphantha nickelsiae
                                                   [Coryphantha sulcata var. nickelsiae]
                                             Coryphantha octacantha
                                             Coryphantha pallida
                                             Coryphantha pseudechinus
                                               Coryphantha pseudechinus subsp. laui
                                                     [Coryphantha laui]
                                               Coryphantha pseudechinus subsp. pseudechinus
                                                     [Coryphantha pusilliflora]
                                             Coryphantha pycnacantha
                                             Coryphantha ramillosa
                                               Coryphantha ramillosa subsp. ramillosa
                                                     [Mammillaria ramillosa]
                                             Coryphantha recurvata
                                                   [Mammillaria recurvata]
                                             Coryphantha retusa
                                             Coryphantha robustispina
                                               Coryphantha robustispina subsp. robustispina
                                                     [Cactus robustispinus]
                                                     [Coryphantha scheeri var. robustispina]
                                                     [Mammillaria robustispina]
                                               Coryphantha robustispina subsp. scheeri
                                                     [Coryphantha muehlenpfordtii]
                                                     [Coryphantha robustispina subsp. uncinata]
                                                     [Coryphantha scheeri var. uncinata]
                                                     [Coryphantha scheeri var. valida]
                                                     [Coryphantha scheeri]
                                                     [Mammillaria scheeri]
                                             Coryphantha salinensis
                                             Coryphantha sulcata
                                     Disocactus
                                             Disocactus nelsonii
                                             Disocactus speciosus
                                               Disocactus speciosus subsp. speciosus
                                                     [Cereus coccineus]
                                     Echinocactus
                                           [Homalocephala]
                                             Echinocactus horizonthalonius
                                               Echinocactus horizonthalonius subsp. horizonthalonius
                                                     [Echinocactus horizontalis]
                                               Echinocactus horizonthalonius subsp. nicholii
                                                     [Echinocactus horizonthalonius var. nicholii]
                                             Echinocactus platyacanthus
                                                   [Echinocactus ingens]
                                             Echinocactus polycephalus
                                               Echinocactus polycephalus subsp. polycephalus
                                                     [Homalocephala polycephala]
                                               Echinocactus polycephalus subsp. xeranthemoides
                                                     [Echinocactus polycephalus var. xeranthemoides]
                                                     [Echinocactus xeranthemoides]
                                                     [Homalocephala polycephala subsp. xeranthemoides]
                                             Echinocactus texensis
                                                   [Homalocephala texensis]
                                     Echinocereus
                                             Echinocereus acifer
                                               Echinocereus acifer subsp. huitcholensis
                                                     [Echinocereus polyacanthus var. huitcholensis]
                                             Echinocereus arizonicus
                                                   [Echinocereus arizonicus subsp. nigrihorridispinus]
                                                   [Echinocereus arizonicus subsp. oldachiorum]
                                                   [Echinocereus coccineus var. arizonicus]
                                                   [Echinocereus triglochidiatus var. arizonicus]
                                             Echinocereus bakeri
                                             Echinocereus bonkerae
                                               Echinocereus bonkerae subsp. apachensis
                                                     [Echinocereus apachensis]
                                               Echinocereus bonkerae subsp. bonkerae
                                                     [Echinocereus fasciculatus subsp. bonkerae]
                                                     [Echinocereus fasciculatus var. bonkerae]
                                                     [Echinocereus fendleri var. bonkerae]
                                             Echinocereus chisoensis
                                             Echinocereus coccineus
                                               Echinocereus coccineus subsp. coccineus
                                                     [Cereus aggregatus]
                                                     [Cereus phoeniceus]
                                                     [Echinocereus aggregatus]
                                                     [Echinocereus canyonensis]
                                                     [Echinocereus coccineus var. inermis]
                                                     [Echinocereus conoideus]
                                                     [Echinocereus engelmannii subsp. decumbens]
                                                     [Echinocereus phoeniceus var. inermis]
                                                     [Echinocereus phoeniceus]
                                                     [Echinocereus triglochidiatus var. inermis]
                                                     [Echinocereus triglochidiatus var. melanacanthus]
                                                     [Echinocereus triglochidiatus var. toroweapensis]
                                               Echinocereus coccineus subsp. gurneyi
                                                     [Echinocereus coccineus var. gurneyi]
                                                     [Echinocereus triglochidiatus var. gurneyi]
                                               Echinocereus coccineus subsp. paucispinus
                                                     [Cereus paucispinus]
                                                     [Echinocereus triglochidiatus var. paucispinus]
                                               Echinocereus coccineus subsp. rosei
                                                     [Echinocereus rosei]
                                               Echinocereus coccineus subsp. santaritensis
                                                     [Echinocereus santaritensis]
                                                     [Mammillaria aggregata]
                                               Echinocereus coccineus subsp. transpecosensis
                                             Echinocereus dasyacanthus
                                               Echinocereus dasyacanthus subsp. ctenoides
                                                     [Cereus ctenoides]
                                                     [Echinocereus ctenoides]
                                               Echinocereus dasyacanthus subsp. dasyacanthus
                                                     [Echinocereus pectinatus var. dasyacanthus]
                                             Echinocereus engelmannii
                                                   [Cereus engelmannii]
                                               Echinocereus engelmannii var. acicularis
                                               Echinocereus engelmannii var. chrysocentrus
                                               Echinocereus engelmannii var. purpureus
                                               Echinocereus engelmannii var. variegatus
                                                     [Cereus engelmannii var. variegatus]
                                                     [Echinocereus engelmannii subsp. variegatus]
                                             Echinocereus enneacanthus
                                               Echinocereus enneacanthus subsp. enneacanthus
                                                     [Cereus dubius]
                                                     [Echinocereus dubius]
                                             Echinocereus fasciculatus
                                               Echinocereus fasciculatus subsp. boyce-thompsonii
                                                     [Echinocereus boyce-thompsonii]
                                                     [Echinocereus fasciculatus var. boyce-thompsonii]
                                                     [Echinocereus fendleri var. boyce-thompsonii]
                                               Echinocereus fasciculatus subsp. fasciculatus
                                                     [Echinocereus abbeae]
                                                     [Echinocereus engelmannii subsp. fasciculatus]
                                                     [Echinocereus fendleri var. fasciculatus]
                                                     [Echinocereus rectispinus var. robustus]
                                             Echinocereus fendleri
                                               Echinocereus fendleri subsp. fendleri
                                                     [Cereus fendleri]
                                                     [Echinocereus hempelii]
                                               Echinocereus fendleri subsp. kuenzleri
                                                     [Echinocereus fendleri var. kuenzleri]
                                                     [Echinocereus kuenzleri]
                                               Echinocereus fendleri subsp. rectispinus
                                                     [Echinocereus fendleri var. rectispinus]
                                                     [Echinocereus rectispinus]
                                             Echinocereus ledingii
                                                   [Echinocereus fendleri var. ledingii]
                                             Echinocereus longisetus
                                             Echinocereus nicholii
                                                   [Echinocereus engelmannii var. nicholii]
                                             Echinocereus pamanesiorum
                                             Echinocereus pectinatus
                                               Echinocereus pectinatus subsp. pectinatus
                                                     [Cereus dasyacanthus var. minor]
                                                     [Cereus pectinatus]
                                             Echinocereus polyacanthus
                                                   [Cereus polyacanthus]
                                                   [Echinocereus triglochidiatus var. polyacanthus]
                                             Echinocereus pseudopectinatus
                                                   [Echinocereus bristolii var. pseudopectinatus]
                                             Echinocereus reichenbachii
                                               Echinocereus reichenbachii subsp. albertii
                                                     [Echinocereus reichenbachii var. albertii]
                                               Echinocereus reichenbachii subsp. armatus
                                               Echinocereus reichenbachii subsp. baileyi
                                                     [Echinocereus reichenbachii var. albispinus]
                                               Echinocereus reichenbachii subsp. perbellus
                                                     [Echinocereus reichenbachii var. perbellus]
                                               Echinocereus reichenbachii subsp. reichenbachii
                                             Echinocereus relictus
                                             Echinocereus rigidissimus
                                               Echinocereus rigidissimus subsp. rigidissimus
                                                     [Echinocereus pectinatus var. rigidissimus]
                                             Echinocereus russanthus
                                                   [Echinocereus chloranthus var. russanthus]
                                             Echinocereus stramineus
                                                   [Cereus stramineus]
                                                   [Echinocereus enneacanthus var. stramineus]
                                             Echinocereus subinermis
                                             Echinocereus triglochidiatus
                                               Echinocereus triglochidiatus subsp. mojavensis
                                                     [Echinocereus mojavensis]
                                                     [Echinocereus triglochidiatus var. mojavensis]
                                               Echinocereus triglochidiatus subsp. triglochidiatus
                                                     [Cereus triglochidiatus]
                                                     [Echinocereus gonacanthus]
                                                     [Echinocereus octacanthus]
                                                     [Echinocereus triglochidiatus var. gonacanthus]
                                             Echinocereus viridiflorus
                                               Echinocereus viridiflorus subsp. chloranthus
                                                     [Cereus chloranthus]
                                                     [Echinocereus chloranthus subsp. rhyolithensis]
                                                     [Echinocereus chloranthus]
                                                     [Echinocereus viridiflorus var. chloranthus]
                                               Echinocereus viridiflorus subsp. cylindricus
                                                     [Echinocereus chloranthus var. cylindricus]
                                                     [Echinocereus viridiflorus var. cylindricus]
                                               Echinocereus viridiflorus subsp. davisii
                                                     [Echinocereus viridiflorus var. davisii]
                                               Echinocereus viridiflorus subsp. viridiflorus
                                                     [Cereus viridiflorus]
                                             Echinocereus yavapaiensis
                                             Echinocereus zapalinamensis
                                             Echinocereus ×kunzei
                                             Echinocereus ×neomexicanus
                                                   [Echinocereus triglochidiatus var. neomexicanus]
                                             Echinocereus ×roetteri
                                                   [Cereus ×roetteri]
                                                   [Echinocereus pectinatus var. neomexicanus]
                                     Echinopsis
                                             Echinopsis rhodacantha
                                                   [Echinocactus coccineus]
                                     Epiphyllum
                                             Epiphyllum hookeri
                                               Epiphyllum hookeri subsp. hookeri
                                                     [Cereus marginatus]
                                             Epiphyllum oxypetalum
                                     Epithelantha
                                             Epithelantha bokei
                                             Epithelantha micromeris
                                                   [Mammillaria micromeris]
                                     Eriosyce
                                             Eriosyce armata
                                                   [Eriosyce curvispina var. armata]
                                             Eriosyce chilensis
                                             Eriosyce iquiquensis
                                             Eriosyce napina
                                             Eriosyce paucicostata
                                               Eriosyce paucicostata subsp. floccosa
                                                     [Pyrrhocactus floccosus]
                                             Eriosyce taltalensis
                                               Eriosyce taltalensis subsp. taltalensis
                                     Ferocactus
                                           [Hamatocactus]
                                             Ferocactus cylindraceus
                                               Ferocactus cylindraceus subsp. cylindraceus
                                                     [Echinocactus acanthodes]
                                               Ferocactus cylindraceus subsp. eastwoodiae
                                                     [Ferocactus acanthodes var. eastwoodiae]
                                                     [Ferocactus cylindraceus var. eastwoodiae]
                                                     [Ferocactus eastwoodiae]
                                               Ferocactus cylindraceus subsp. lecontei
                                                     [Echinocactus lecontei]
                                                     [Ferocactus acanthodes var. lecontei]
                                                     [Ferocactus cylindraceus var. lecontei]
                                                     [Ferocactus lecontei]
                                             Ferocactus diguetii
                                             Ferocactus emoryi
                                               Ferocactus emoryi subsp. covillei
                                               Ferocactus emoryi subsp. emoryi
                                                     [Echinocactus emoryi]
                                             Ferocactus latispinus
                                             Ferocactus uncinatus
                                                   [Ancistrocactus uncinatus var. wrightii]
                                                   [Ancistrocactus uncinatus]
                                                   [Glandulicactus uncinatus]
                                                   [Hamatocactus uncinatus]
                                                   [Sclerocactus uncinatus var. wrightii]
                                                   [Sclerocactus uncinatus]
                                             Ferocactus viridescens
                                               Ferocactus viridescens subsp. viridescens
                                                     [Ferocactus acanthodes]
                                             Ferocactus wislizeni
                                               Ferocactus wislizeni subsp. wislizeni
                                                     [Echinocactus wislizeni]
                                                     [Ferocactus wislizeni subsp. ajoensis]
                                                     [Ferocactus wislizeni var. ajoensis]
                                     Harrisia
                                             Harrisia fragrans
                                               Harrisia fragrans subsp. aboriginum
                                                     [Harrisia aboriginum]
                                               Harrisia fragrans subsp. fragrans
                                                     [Cereus eriophorus var. fragrans]
                                                     [Cereus fragrans]
                                             Harrisia portoricensis
                                     Leptocereus
                                             Leptocereus grantianus
                                     Leuchtenbergia
                                             Leuchtenbergia principis
                                     Lophophora
                                             Lophophora williamsii
                                     Mammillaria
                                             Mammillaria elongata
                                             Mammillaria geminispina
                                               Mammillaria geminispina subsp. geminispina
                                                     [Mammillaria elegans]
                                             Mammillaria heyderi
                                               Mammillaria heyderi subsp. gummifera
                                                     [Mammillaria gummifera]
                                               Mammillaria heyderi subsp. hemisphaerica
                                                     [Mammillaria gummifera var. applanata]
                                                     [Mammillaria gummifera var. hemisphaerica]
                                                     [Mammillaria heyderi var. applanata]
                                                     [Mammillaria heyderi var. hemisphaerica]
                                               Mammillaria heyderi subsp. heyderi
                                                     [Mammillaria gummifera var. macdougalii]
                                                     [Mammillaria heyderi subsp. macdougalii]
                                                     [Mammillaria heyderi var. bullingtoniana]
                                                     [Mammillaria heyderi var. macdougalii]
                                                     [Mammillaria macdougalii]
                                                     [Neomammillaria macdougalii]
                                               Mammillaria heyderi subsp. meiacantha
                                                     [Mammillaria gummifera var. meiacantha]
                                                     [Mammillaria heyderi var. meiacantha]
                                                     [Mammillaria meiacantha]
                                             Mammillaria johnstonii
                                             Mammillaria lasiacantha
                                               Mammillaria lasiacantha subsp. lasiacantha
                                             Mammillaria longimamma
                                             Mammillaria monochrysacantha
                                             Mammillaria plumosa
                                             Mammillaria pottsii
                                                   [Coryphantha pottsiana]
                                             Mammillaria rhodantha
                                               Mammillaria rhodantha subsp. mccartenii
                                             Mammillaria sphaerica
                                                   [Mammillaria longimamma var. sphaerica]
                                             Mammillaria wagneriana
                                     Marginatocereus
                                             Marginatocereus marginatus
                                                   [Pachycereus marginatus]
                                     Myrtillocactus
                                             Myrtillocactus cochal
                                             Myrtillocactus geometrizans
                                     Pachycereus
                                           [Lophocereus]
                                             Pachycereus pecten-aboriginum
                                             Pachycereus schottii
                                                   [Cereus schottii]
                                                   [Lophocereus schottii]
                                     Parodia
                                             Parodia microsperma
                                     Parrycactus
                                             Parrycactus echidne
                                                   [Ferocactus echidne]
                                     Pediocactus
                                           [Navajoa]
                                           [Utahia]
                                             Pediocactus bradyi
                                               Pediocactus bradyi subsp. bradyi
                                                     [Toumeya bradyi]
                                               Pediocactus bradyi subsp. despainii
                                                     [Pediocactus despainii]
                                               Pediocactus bradyi subsp. winkleri
                                                     [Pediocactus winkleri]
                                             Pediocactus knowltonii
                                             Pediocactus paradinei
                                             Pediocactus peeblesianus
                                               Pediocactus peeblesianus subsp. peeblesianus
                                                     [Echinocactus peeblesianus]
                                                     [Navajoa peeblesiana]
                                                     [Pediocactus peeblesianus var. maianus]
                                             Pediocactus sileri
                                                   [Echinocactus sileri]
                                                   [Utahia sileri]
                                             Pediocactus simpsonii
                                                   [Echinocactus simpsonii]
                                                   [Pediocactus hermannii]
                                                   [Pediocactus simpsonii var. minor]
                                     Pelecyphora
                                           [Encephalocarpus]
                                           [Neobesseya]
                                             Pelecyphora alversonii
                                                   [Coryphantha alversonii]
                                                   [Escobaria alversonii]
                                                   [Mammillaria alversonii]
                                             Pelecyphora chihuahuensis
                                               Pelecyphora chihuahuensis subsp. chihuahuensis
                                                     [Coryphantha strobiliformis]
                                                     [Escobaria strobiliformis]
                                             Pelecyphora dasyacantha
                                               Pelecyphora dasyacantha subsp. chaffeyi
                                                     [Coryphantha chaffeyi]
                                               Pelecyphora dasyacantha subsp. dasyacantha
                                                     [Coryphantha chlorantha]
                                                     [Coryphantha dasyacantha]
                                                     [Escobaria dasyacantha]
                                                     [Mammillaria chlorantha]
                                             Pelecyphora duncanii
                                                   [Coryphantha duncanii]
                                                   [Escobaria dasyacantha var. duncanii]
                                                   [Escobaria duncanii]
                                             Pelecyphora hesteri
                                               Pelecyphora hesteri subsp. hesteri
                                                     [Coryphantha hesteri]
                                                     [Escobaria hesteri]
                                             Pelecyphora laredoi
                                                   [Coryphantha laredoi]
                                             Pelecyphora macromeris
                                               Pelecyphora macromeris subsp. macromeris
                                                     [Coryphantha macromeris]
                                                     [Mammillaria macromeris]
                                               Pelecyphora macromeris subsp. runyonii
                                                     [Coryphantha macromeris subsp. runyonii]
                                                     [Coryphantha macromeris var. runyonii]
                                                     [Coryphantha runyonii]
                                             Pelecyphora minima
                                                   [Coryphantha minima]
                                                   [Coryphantha nellieae]
                                                   [Escobaria minima]
                                                   [Escobaria nellieae]
                                                   [Mammillaria nellieae]
                                             Pelecyphora missouriensis
                                               Pelecyphora missouriensis subsp. missouriensis
                                                     [Coryphantha missouriensis var. caespitosa]
                                                     [Coryphantha missouriensis var. marstonii]
                                                     [Coryphantha missouriensis var. robustior]
                                                     [Coryphantha missouriensis]
                                                     [Coryphantha similis]
                                                     [Escobaria missouriensis subsp. navajoensis]
                                                     [Escobaria missouriensis var. caespitosa]
                                                     [Escobaria missouriensis var. marstonii]
                                                     [Escobaria missouriensis var. robustior]
                                                     [Escobaria missouriensis var. similis]
                                                     [Escobaria missouriensis]
                                                     [Mammillaria missouriensis]
                                                     [Mammillaria similis var. robustior]
                                                     [Mammillaria similis]
                                                     [Neobesseya missouriensis]
                                                     [Neobesseya wissmannii]
                                             Pelecyphora robbinsiorum
                                                   [Coryphantha robbinsiorum]
                                                   [Escobaria robbinsiorum]
                                             Pelecyphora sneedii
                                               Pelecyphora sneedii subsp. orcuttii
                                                     [Coryphantha orcuttii]
                                                     [Coryphantha organensis]
                                                     [Coryphantha sneedii var. orcuttii]
                                                     [Coryphantha sneedii var. organensis]
                                                     [Coryphantha sneedii var. sandbergii]
                                                     [Coryphantha sneedii var. villardii]
                                                     [Coryphantha strobiliformis var. orcuttii]
                                                     [Escobaria orcuttii var. koenigii]
                                                     [Escobaria orcuttii]
                                                     [Escobaria organensis]
                                                     [Escobaria sandbergii]
                                                     [Escobaria villardii]
                                                     [Neolloydia orcuttii]
                                               Pelecyphora sneedii subsp. sneedii
                                                     [Coryphantha albicolumnaria]
                                                     [Coryphantha sneedii var. guadalupensis]
                                                     [Coryphantha sneedii var. leei]
                                                     [Coryphantha sneedii]
                                                     [Escobaria albicolumnaria]
                                                     [Escobaria guadalupensis]
                                                     [Escobaria leei]
                                                     [Escobaria sneedii subsp. leei]
                                                     [Escobaria sneedii var. leei]
                                                     [Escobaria sneedii]
                                             Pelecyphora strobiliformis
                                                   [Encephalocarpus strobiliformis]
                                             Pelecyphora tuberculosa
                                                   [Coryphantha dasyacantha var. varicolor]
                                                   [Coryphantha strobiliformis var. durispina]
                                                   [Coryphantha tuberculosa]
                                                   [Escobaria dasyacantha var. varicolor]
                                                   [Escobaria tuberculosa var. varicolor]
                                                   [Escobaria tuberculosa]
                                                   [Mammillaria tuberculosa]
                                             Pelecyphora vivipara
                                                   [Cactus viviparus]
                                                   [Coryphantha deserti]
                                                   [Coryphantha fragrans]
                                                   [Coryphantha rosea]
                                                   [Coryphantha vivipara var. arizonica]
                                                   [Coryphantha vivipara var. bisbeeana]
                                                   [Coryphantha vivipara var. buoflama]
                                                   [Coryphantha vivipara var. deserti]
                                                   [Coryphantha vivipara var. kaibabensis]
                                                   [Coryphantha vivipara var. neomexicana]
                                                   [Coryphantha vivipara var. radiosa]
                                                   [Coryphantha vivipara var. rosea]
                                                   [Coryphantha vivipara]
                                                   [Escobaria vivipara var. arizonica]
                                                   [Escobaria vivipara var. bisbeeana]
                                                   [Escobaria vivipara var. deserti]
                                                   [Escobaria vivipara var. neomexicana]
                                                   [Escobaria vivipara var. radiosa]
                                                   [Escobaria vivipara var. rosea]
                                                   [Escobaria vivipara]
                                                   [Mammillaria deserti]
                                                   [Mammillaria radiosa]
                                                   [Mammillaria vivipara var. arizonica]
                                                   [Mammillaria vivipara]
                                     Peniocereus
                                             Peniocereus greggii
                                               Peniocereus greggii subsp. greggii
                                                     [Cereus greggii]
                                               Peniocereus greggii subsp. transmontanus
                                                     [Cereus greggii var. transmontanus]
                                                     [Peniocereus greggii var. transmontanus]
                                             Peniocereus striatus
                                     Pilosocereus
                                             Pilosocereus leucocephalus
                                             Pilosocereus robinii
                                     Schlumbergera
                                     Sclerocactus
                                           [Ancistrocactus]
                                           [Glandulicactus]
                                             Sclerocactus brevihamatus
                                               Sclerocactus brevihamatus subsp. tobuschii
                                             Sclerocactus brevispinus
                                                   [Sclerocactus whipplei var. ilseae]
                                             Sclerocactus cloverae
                                                   [Sclerocactus cloverae subsp. brackii]
                                                   [Sclerocactus whipplei var. heilii]
                                                   [Sclerocactus whipplei var. reevesii]
                                             Sclerocactus glaucus
                                                   [Echinocactus glaucus]
                                                   [Sclerocactus whipplei var. glaucus]
                                             Sclerocactus intertextus
                                                   [Echinocactus intertextus]
                                                   [Echinomastus intertextus subsp. dasyacanthus]
                                                   [Echinomastus intertextus var. dasyacanthus]
                                                   [Echinomastus intertextus]
                                                   [Neolloydia intertexta var. dasyacantha]
                                                   [Neolloydia intertexta]
                                                   [Sclerocactus intertextus var. dasyacanthus]
                                             Sclerocactus johnsonii
                                               Sclerocactus johnsonii subsp. acunensis
                                                     [Echinomastus erectocentrus var. acunensis]
                                                     [Neolloydia erectocentra var. acunensis]
                                               Sclerocactus johnsonii subsp. erectocentrus
                                                     [Echinomastus erectocentrus]
                                                     [Neolloydia erectocentra]
                                                     [Sclerocactus erectocentrus]
                                               Sclerocactus johnsonii subsp. johnsonii
                                                     [Echinocactus johnsonii var. lutescens]
                                                     [Echinocactus johnsonii]
                                                     [Echinomastus johnsonii var. lutescens]
                                                     [Echinomastus johnsonii]
                                                     [Neolloydia johnsonii]
                                             Sclerocactus mariposensis
                                                   [Echinomastus mariposensis]
                                             Sclerocactus mesae-verdae
                                               Sclerocactus mesae-verdae subsp. depressus
                                               Sclerocactus mesae-verdae subsp. mesae-verdae
                                                     [Coloradoa mesae-verdae]
                                                     [Echinocactus mesae-verdae]
                                                     [Pediocactus mesae-verdae]
                                             Sclerocactus nyensis
                                             Sclerocactus papyracanthus
                                                   [Echinocactus papyracanthus]
                                                   [Mammillaria papyracantha]
                                                   [Pediocactus papyracanthus]
                                                   [Toumeya papyracantha]
                                             Sclerocactus parviflorus
                                               Sclerocactus parviflorus subsp. intermedius
                                                     [Sclerocactus intermedius]
                                                     [Sclerocactus parviflorus var. intermedius]
                                               Sclerocactus parviflorus subsp. parviflorus
                                                     [Sclerocactus contortus]
                                               Sclerocactus parviflorus subsp. terrae-canyonae
                                                     [Sclerocactus terrae-canyonae]
                                             Sclerocactus polyancistrus
                                             Sclerocactus pubispinus
                                             Sclerocactus sileri
                                                   [Sclerocactus pubispinus var. sileri]
                                             Sclerocactus spinosior
                                                   [Echinocactus whipplei var. spinosior]
                                                   [Sclerocactus pubispinus var. spinosior]
                                             Sclerocactus wetlandicus
                                             Sclerocactus whipplei
                                               Sclerocactus whipplei subsp. whipplei
                                                     [Echinocactus whipplei]
                                                     [Pediocactus whipplei]
                                             Sclerocactus wrightiae
                                     Selenicereus
                                             Selenicereus grandiflorus
                                               Selenicereus grandiflorus subsp. hondurensis
                                                     [Selenicereus hondurensis]
                                             Selenicereus pteranthus
                                             Selenicereus undatus
                                                   [Hylocereus undatus]
                                     Soehrensia
                                             Soehrensia thelegona
                                                   [Trichocereus thelegonus]
                                     Stenocactus
                                             Stenocactus multicostatus
                                                   [Echinofossulocactus multicostatus]
                                             Stenocactus ×irregularis
                                     Stenocereus
                                             Stenocereus alamosensis
                                             Stenocereus thurberi
                                               Stenocereus thurberi subsp. thurberi
                                                     [Cereus thurberi]
                                                     [Lemaireocereus thurberi]
                                     Thelocactus
                                             Thelocactus bicolor
                                               Thelocactus bicolor subsp. heterochromus
                                                     [Thelocactus heterochromus]
                                             Thelocactus leucacanthus
                                     Trichocereus
                                     Turbinicarpus
                                           [Toumeya]
                                             Turbinicarpus schmiedickeanus
                                               Turbinicarpus schmiedickeanus subsp. klinkerianus
                                                     [Turbinicarpus klinkerianus]
                                     Weberocereus
                                             Weberocereus tunilla
                                               Weberocereus tunilla subsp. tunilla
                                                     [Cereus gonzalezii]
                              Opuntioideae
                                     Austrocylindropuntia
                                             Austrocylindropuntia subulata
                                               Austrocylindropuntia subulata subsp. subulata
                                                     [Opuntia subulata]
                                     Brasiliopuntia
                                             Brasiliopuntia brasiliensis
                                     Consolea
                                             Consolea corallicola
                                             Consolea macracantha
                                                   [Opuntia macracantha]
                                     Cylindropuntia
                                             Cylindropuntia abyssi
                                                   [Opuntia abyssi]
                                             Cylindropuntia acanthocarpa
                                               Cylindropuntia acanthocarpa subsp. acanthocarpa
                                                     [Opuntia acanthocarpa]
                                               Cylindropuntia acanthocarpa subsp. coloradensis
                                                     [Cylindropuntia acanthocarpa var. coloradensis]
                                                     [Opuntia acanthocarpa var. coloradensis]
                                               Cylindropuntia acanthocarpa subsp. major
                                                     [Cylindropuntia acanthocarpa subsp. ramosa]
                                                     [Cylindropuntia acanthocarpa var. major]
                                                     [Cylindropuntia acanthocarpa var. ramosa]
                                                     [Opuntia acanthocarpa var. major]
                                                     [Opuntia acanthocarpa var. ramosa]
                                               Cylindropuntia acanthocarpa var. thornberi
                                                     [Cylindropuntia acanthocarpa subsp. thornberi]
                                                     [Opuntia acanthocarpa var. thornberi]
                                                     [Opuntia thornberi]
                                             Cylindropuntia alcahes
                                               Cylindropuntia alcahes var. burrageana
                                             Cylindropuntia arbuscula
                                                   [Opuntia arbuscula]
                                             Cylindropuntia bernardina
                                                   [Opuntia californica var. parkeri]
                                             Cylindropuntia bigelowii
                                               Cylindropuntia bigelowii subsp. bigelowii
                                                     [Opuntia bigelovii]
                                               Cylindropuntia bigelowii subsp. ciribe
                                                     [Cylindropuntia bigelovii var. ciribe]
                                             Cylindropuntia californica
                                               Cylindropuntia californica subsp. californica
                                                     [Opuntia californica]
                                               Cylindropuntia californica subsp. delgadilloana
                                                     [Cylindropuntia delgadilloana]
                                               Cylindropuntia californica subsp. rosarica
                                                     [Cylindropuntia californica var. rosarica]
                                             Cylindropuntia cholla
                                             Cylindropuntia davisii
                                                   [Opuntia davisii]
                                                   [Opuntia tunicata var. davisii]
                                             Cylindropuntia echinocarpa
                                                   [Opuntia echinocarpa]
                                                   [Opuntia wigginsii]
                                             Cylindropuntia fulgida
                                                   [Opuntia fulgida]
                                               Cylindropuntia fulgida var. mamillata
                                                     [Opuntia fulgida var. mamillata]
                                             Cylindropuntia imbricata
                                               Cylindropuntia imbricata subsp. imbricata
                                                     [Opuntia arborescens]
                                                     [Opuntia imbricata]
                                               Cylindropuntia imbricata subsp. spinosior
                                                     [Cylindropuntia imbricata var. spinosior]
                                                     [Cylindropuntia spinosior]
                                                     [Opuntia spinosior var. neomexicana]
                                                     [Opuntia spinosior]
                                             Cylindropuntia kleiniae
                                                   [Opuntia kleiniae]
                                             Cylindropuntia leptocaulis
                                                   [Opuntia leptocaulis]
                                             Cylindropuntia molesta
                                               Cylindropuntia molesta subsp. clavellina
                                                     [Cylindropuntia molesta var. clavellina]
                                               Cylindropuntia molesta subsp. molesta
                                             Cylindropuntia prolifera
                                             Cylindropuntia ramosissima
                                                   [Opuntia ramosissima]
                                             Cylindropuntia tesajo
                                                   [Opuntia tesajo]
                                             Cylindropuntia thurberi
                                               Cylindropuntia thurberi subsp. versicolor
                                                     [Cylindropuntia versicolor]
                                                     [Opuntia versicolor]
                                             Cylindropuntia tunicata
                                                   [Opuntia tunicata]
                                             Cylindropuntia whipplei
                                               Cylindropuntia whipplei subsp. enodis
                                                     [Cylindropuntia whipplei var. enodis]
                                             Cylindropuntia ×congesta
                                             Cylindropuntia ×deserta
                                             Cylindropuntia ×fosbergii
                                                   [Cylindropuntia bigelovii]
                                             Cylindropuntia ×kelvinensis
                                                   [Opuntia ×kelvinensis]
                                             Cylindropuntia ×multigeniculata
                                                   [Opuntia whipplei var. multigeniculata]
                                             Cylindropuntia ×neoarbuscula
                                                   [Opuntia ×neoarbuscula]
                                             Cylindropuntia ×tetracantha
                                                   [Opuntia ×tetracantha]
                                             Cylindropuntia ×viridiflora
                                                   [Opuntia ×viridiflora]
                                             Cylindropuntia ×vivipara
                                                   [Opuntia ×vivipara]
                                     Grusonia
                                           [Corynopuntia]
                                             Grusonia aggeria
                                                   [Opuntia aggeria]
                                             Grusonia clavata
                                                   [Corynopuntia clavata]
                                                   [Opuntia clavata]
                                             Grusonia emoryi
                                                   [Corynopuntia emoryi]
                                                   [Grusonia stanlyi]
                                                   [Opuntia emoryi]
                                                   [Opuntia stanlyi var. peeblesiana]
                                                   [Opuntia stanlyi]
                                             Grusonia grahamii
                                                   [Corynopuntia grahamii]
                                             Grusonia invicta
                                             Grusonia kunzei
                                                   [Corynopuntia kunzei]
                                                   [Grusonia wrightiana]
                                                   [Opuntia kunzei]
                                             Grusonia marenae
                                             Grusonia parishiorum
                                             Grusonia schottii
                                                   [Opuntia densispina]
                                     Maihueniopsis
                                             Maihueniopsis glomerata
                                     Micropuntia
                                             Micropuntia pulchella
                                                   [Grusonia pulchella]
                                                   [Opuntia pulchella]
                                     Opuntia
                                           [Nopalea]
                                             Opuntia aciculata
                                             Opuntia atrispina
                                             Opuntia auberi
                                                   [Nopalea auberi]
                                             Opuntia aurea
                                                   [Opuntia erinacea var. aurea]
                                             Opuntia basilaris
                                                   [Opuntia basilaris var. heilii]
                                               Opuntia basilaris var. longiareolata
                                               Opuntia basilaris var. treleasei
                                                     [Opuntia treleasei]
                                             Opuntia chaffeyi
                                             Opuntia chlorotica
                                             Opuntia cochenillifera
                                                   [Nopalea cochenillifera]
                                             Opuntia decumbens
                                             Opuntia dejecta
                                                   [Nopalea dejecta]
                                             Opuntia engelmannii
                                               Opuntia engelmannii subsp. engelmannii
                                                     [Opuntia caesia]
                                                     [Opuntia discata]
                                                     [Opuntia engelmannii var. flavescens]
                                                     [Opuntia flavescens]
                                                     [Opuntia microcarpa]
                                                     [Opuntia phaeacantha var. discata]
                                                     [Opuntia valida]
                                               Opuntia engelmannii var. cacanapa
                                                     [Opuntia ellisiana]
                                               Opuntia engelmannii var. dulcis
                                                     [Opuntia dulcis]
                                               Opuntia engelmannii var. flavispina
                                                     [Opuntia phaeacantha var. flavispina]
                                               Opuntia engelmannii var. laevis
                                                     [Opuntia laevis]
                                                     [Opuntia phaeacantha var. laevis]
                                               Opuntia engelmannii var. lindheimeri
                                                     [Opuntia lindheimeri var. subarmata]
                                                     [Opuntia lindheimeri]
                                               Opuntia engelmannii var. linguiformis
                                                     [Opuntia lindheimeri var. linguiformis]
                                                     [Opuntia linguiformis]
                                             Opuntia ficus-indica
                                                   [Opuntia castillae]
                                             Opuntia fragilis
                                                   [Opuntia brachyarthra]
                                                   [Opuntia fragilis var. brachyarthra]
                                                   [Opuntia fragilis var. denudata]
                                             Opuntia gosseliniana
                                                   [Opuntia macrocentra var. santa-rita]
                                                   [Opuntia santa-rita]
                                                   [Opuntia violacea var. gosseliniana]
                                                   [Opuntia violacea var. santa-rita]
                                             Opuntia hondurensis
                                             Opuntia humifusa
                                               Opuntia humifusa subsp. humifusa
                                                     [Opuntia compressa]
                                               Opuntia humifusa subsp. minor
                                                     [Opuntia rafinesquei]
                                             Opuntia leucotricha
                                             Opuntia littoralis
                                             Opuntia macrocentra
                                                   [Opuntia violacea var. macrocentra]
                                                   [Opuntia violacea]
                                             Opuntia macrorhiza
                                                   [Opuntia compressa var. macrorhiza]
                                                   [Opuntia greenei]
                                                   [Opuntia mesacantha var. greenii]
                                                   [Opuntia plumbea]
                                                   [Opuntia stenochila]
                                                   [Opuntia tortispina]
                                             Opuntia microdasys
                                             Opuntia orbiculata
                                                   [Opuntia dillei]
                                             Opuntia phaeacantha
                                                   [Opuntia angustata]
                                                   [Opuntia engelmannii var. cyclodes]
                                                   [Opuntia eocarpa]
                                                   [Opuntia recurvospina]
                                                   [Opuntia rubrifolia]
                                                   [Opuntia zuniensis]
                                               Opuntia phaeacantha var. camanchica
                                                     [Opuntia phaeacantha var. brunnea]
                                               Opuntia phaeacantha var. castorea
                                               Opuntia phaeacantha var. chihuahuensis
                                                     [Opuntia chihuahuensis]
                                               Opuntia phaeacantha var. major
                                                     [Opuntia arizonica]
                                                     [Opuntia blakeana]
                                                     [Opuntia gilvescens]
                                                     [Opuntia toumeyi]
                                               Opuntia phaeacantha var. superbospina
                                                     [Opuntia superbospina]
                                             Opuntia pinkavae
                                                   [Opuntia basilaris var. woodburyi]
                                             Opuntia polyacantha
                                                   [Opuntia erinacea var. utahensis]
                                                   [Opuntia heacockiae]
                                                   [Opuntia juniperina]
                                                   [Opuntia missouriensis]
                                                   [Opuntia polyacantha var. juniperina]
                                                   [Opuntia polyacantha var. rufispina]
                                                   [Opuntia polyacantha var. watsonii]
                                                   [Opuntia rutila]
                                                   [Opuntia schweriniana]
                                                   [Opuntia sphaerocarpa]
                                                   [Opuntia spirocentra]
                                               Opuntia polyacantha var. arenaria
                                                     [Opuntia arenaria]
                                                     [Opuntia polyacantha subsp. arenaria]
                                               Opuntia polyacantha var. erinacea
                                                     [Opuntia diploursina]
                                                     [Opuntia erinacea var. ursina]
                                                     [Opuntia erinacea]
                                                     [Opuntia polyacantha subsp. erinacea]
                                                     [Opuntia ursina]
                                               Opuntia polyacantha var. hystricina
                                                     [Opuntia erinacea var. hystricina]
                                                     [Opuntia hystricina]
                                                     [Opuntia rhodantha var. spinosior]
                                                     [Opuntia rhodantha]
                                               Opuntia polyacantha var. nicholii
                                                     [Opuntia erinacea var. xanthostemma]
                                                     [Opuntia nicholii]
                                               Opuntia polyacantha var. trichophora
                                                     [Opuntia missouriensis var. trichophora]
                                                     [Opuntia trichophora]
                                             Opuntia pottsii
                                                   [Opuntia ballii]
                                                   [Opuntia delicata]
                                                   [Opuntia filipendula]
                                                   [Opuntia macrorhiza var. pottsii]
                                                   [Opuntia tenuispina]
                                             Opuntia pycnantha
                                             Opuntia robusta
                                             Opuntia rufida
                                             Opuntia schumannii
                                             Opuntia stricta
                                                   [Opuntia stricta subsp. esparzae]
                                             Opuntia strigil
                                             Opuntia sulphurea
                                                   [Opuntia albisetosa]
                                             Opuntia triacanthos
                                                   [Opuntia abjecta]
                                             Opuntia tuna
                                             Opuntia ×curvispina
                                                   [Opuntia littoralis var. martiniana]
                                                   [Opuntia ×martiniana]
                                             Opuntia ×cymochila
                                             Opuntia ×occidentalis
                                             Opuntia ×vaseyi
                                                   [Opuntia ×rugosa]
                                             Opuntia ×wootonii
                                                   [Opuntia phaeacantha var. wootonii]
                                     Pereskiopsis
                                             Pereskiopsis kellermanii
                                             Pereskiopsis porteri
                                     Salmonopuntia
                                             Salmonopuntia salmiana
                                                   [Opuntia salmiana]
                                     Tephrocactus
                                             Tephrocactus alexanderi
                                             Tephrocactus articulatus
                              Pereskioideae
                                     Pereskia
                                             Pereskia bahiensis
                                             Pereskia nemorosa
                                             Pereskia sacharosa